5.7.1 smtp;550 Unable to relay

QUESTION 
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One of our email addresses is encountering problems with receiving email. The sender gets a message: 

The following recipient(s) could not be reached: 

User1@domain.com on 11/01/00 8:14 AM 

The originator does not have permission to submit message 
dns;Sendingdomain.com failed 5.7.1 smtp;550 
5.7.1 Unable to relay for User1@domain.com 

ANSWER 
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The 550 5.7.1 error has to do with the sender not able to relay mail from their server to the recipient. This is a server configuration problem on their end. Please see the Microsoft Knowledge Base article below. 

http://support.microsoft.com/default.aspx?scid=kb;en-us;q289553
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